Nate Hardy

Nate Hardy

Assistant Professor

Lab Page

321 Funchess Hall
+1 334-844-1171 (Office)

Areas of Interest:

Insect evolution and systematics, species diversification of plant-feeding insect, phylogenetics.


2008 Ph.D., University of California, Davis (Entomology)
2002 B.S., Cornell University (Entomology)

Professional Appointments:

2013 Assistant Professor, Auburn University
2011-2013 Postdoctoral Researcher, University of New Mexico / Cleveland Museum of Natural History
2008-2011 Research Entomologist, Queensland Department of Primary Industries and Fisheries (Australia)
2008 Postdoctoral Researcher, University of California, Davis


  • Insect Systematics (ENTM 5300/6300)
  • Insect Morphology (ENTM 5440/6440)
Recent Research Publications:
  1. Ross, L., Gardner, A., Hardy, N., & West, S. A. (2013). Ecology, Not the Genetics of Sex Determination, Determines Who Helps in Eusocial Populations. Current Biology.
  2. Miller, K., Jean, A., Alarie, Y., Hardy, N., & Gibson, R. (2013). Phylogenetic placement of North American subterranean diving beetles (Insecta: Coleoptera: Dytiscidae). Arthropod Systematics and Phylogeny 71: 75-90.
  3. Hodgson CJ, Hardy NB. 2013. The phylogeny of the superfamily Coccoidea (Hemiptera: Sternorrhyncha) based on the morphology of extant and extinct macropterous males. Systematic Entomology 38, 794-804.
  4. Hardy NB. 2013. The status and future of scale insects (Coccoidea) systematics. Systematic Entomology 38, 453-458.
  5. Hardy NB, Cook LG. 2012. Testing for ecological limitation of diversification: a case study using parasitic plants. The American Naturalist 180, 438-449.
  6. Ross L, Hardy NB, Okusu A, Normark BB. 2012. Large population size predicts the distribution of asexuality in scale insects. Evolution 67, 196-206.
  7. Cranston PS, Hardy NB, Morse GE. 2012. A dated molecular phylogeny for the Chironomidae (Diptera). Systematic Entomology 37, 172-188.
  8. Hardy NB, Henderson RC. 2011. Revision of Poliaspis (Hemiptera: Coccoidea: Diaspididae), with descriptions of eight new species from Australia. Zookeys 137, 1-40.
  9. Hardy NB, Beardsley JW, Gullan PJ. 2011. Uncovering diversity of Australian Eucalyptus-constrained felt scales (Hemiptera: Coccoidea: Eriococcidae). Systematic Entomology 36, 497-528.
  10. Gruwell ME, Hardy NB, Gullan PJ, Dittmar K. 2010. Evolutionary relationships among primary endosymbionts in the mealybug subfamily Phenacoccinae (Hemiptera: Coccoidea: Pseudococcidae). Applied and Environmental Microbiology 76, 7521-7525.
  11. Hardy NB, Gullan PJ. 2010. Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea, Eriococcidae) and descriptions of a new genus and nine new species. Zookeys 58, 1-74.
  12. Cook LG, Edwards RD, Crisp M, Hardy NB. 2010. Need morphology always be required for new species descriptions? Invertebrate Systematics 24, 322-326.
  13. Hardy NB, Cook LG. 2010. Gall-induction in insects: evolutionary dead-end or speciation driver? BMC Evolutionary Biology 10, 257.
  14. Cranston PS, Hardy NB, Morse GE, Puslednick L, McCluen SR. 2010. When molecules and morphology concur: the ‘gondwanan’ midges (Diptera: Chironomidae). Systematic Entomology 35, 636-648.
  15. Winterton SL, Hardy NB, Wiegman BM. 2010. On wings of lace: phylogeny and Bayesian divergence time estimates of Neuropterida (Insecta) based on morphological and molecular data. Systematic Entomology 35, 349-378.
  16. Gullan PJ, Kaydan MB, Hardy NB. 2010. Molecular phylogeny and species recognition in the mealybug genus Ferrisia (Hemiptera: Pseudococcidae). Systematic Entomology 35, 329-339.
  17. Hardy NB, Gullan, PJ. 2009. Opisthoscelis nigra Froggatt (Eriococcidae) is an armoured scale Maskellia nigra (Froggatt) (Diaspididae). pp. 59–62. In: Branco, M., Franco, J.C. & Hodgson, C. (Eds) Proceedings of the XI International Symposium of Scale Insect Studies, Oeiras, Portugal, 24–27 September 2007. ISA Press, Lisbon, Portugal.
  18. Kondo T, Hardy NB. 2008. Redesciption of Inglesia vitrea Cockerell (Hemiptera, Coccidae) and its transfer to the genus Pseudokermes Cockerell. Zookeys 3, 11-21.
  19. Hardy NB, Gullan PJ, Henderson RC, Cook LG. 2008. Relationships among scale insects (Hemiptera: Coccoidea: Eriococcidae) of southern beech, Nothofagus (Nothofagaceae), with the first descriptions of Australian species of the Nothofagus-feeding genus Madarococcus Hoy. Invertebrate Systematics 22, 365-405.
  20. Kondo T, Hardy NB. 2008. Synonymy of Etiennea Matile-Ferrero with Hemilecanium Newstead (Hemiptera: Coccidae), based on morphology of adult females, adult males and first-instar nymphs, and description of a new species from Okinawa, Japan. Entomological Science 11, 189-213.
  21. Hardy NB, Gullan PJ. 2008. Pseudococcini Cockerell, 1905: proposed precedence over Sphaerococcini Cockerell, 1899. Case number 3424. The Bulletin of Zoological Nomenclature 65, 24-64.
  22. Hardy NB, Gullan PJ, Hodgson CJ. 2008. A classification of mealybugs (Hemiptera: Pseudococcidae) based on integrated molecular and morphological data. Systematic Entomology 33, 51-71.
  23. Hardy NB. 2007. A new mealybug species (Hemiptera: Coccoidea: Pseudococcidae) from the ant-plant Hydnophytum (Rubiaceae) in Fiji. Fiji Arthropods In: Evenhuis NL & Bickel DJ (eds). Fiji Arthropods IX. Bishop Museum Occasional Papers. 94, 35-40.
  24. Hardy NB, Gullan PJ. 2007. A new species of Phacelococcus Miller (Hemiptera: Coccoidea: Eriococcidae) on Bursaria (Pittosporaceae). The Australian Entomologist 34, 85-91.
  25. Hardy NB. 2007. Phylogenetic utility of dynamin and triose phosphate isomerase. Systematic Entomology 32, 396-403.
  26. Hardy NB, Gullan PJ. 2007. A new genus and four new species of felt scales on Eucalyptus (Hemiptera: Coccoidea: Eriococcidae) in south-eastern Australia. Australian Journal of Entomology 46, 106-120.
  27. Kondo T, Hardy NB, Cook LG, Gullan PJ. 2006. Description of two new genera and species of Eriococcidae (Hemiptera: Coccoidea) from southern South America. Zootaxa 1349, 19-36.

Last Updated: April 21, 2014

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