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 Areas of Interest:
Research in systematics of the weevils of the family Curculionidae (Coleoptera), especially the members of the tribe Anthonomini. 
Education:
B.S. 1968; Brigham Young University.
M.S. 1970, Zoology;  Brigham Young University.
Ph.D 1975, Entomology; Texas A&M University.  
Teaching Responsibilities:
Insects (ENTM 2040)
General Entomology (ENTM 3040)
 Entomology for Educators (ENTM 6010)
 Insect Morphology (ENTM 4220 and 7220)
 
Professional Responsibilities:
 
Departmental Graduate Program Officer
 
Other Relevant Achievements and Positions Held:
 
1987- The Coleopterists Bulletin, Outstanding Paper Award.
Biosystematics and Ecology of the Boll Weevil research team, investigation of systematic and ecological relationships of the boll weevil and related weevils.
1981-1985 - Field work in Brazil, Colombia, the Dominican Republic, Guatemala, Honduras, Mexico, Peru and Venezuela.
1993 - Taxonomy of anthonomine weevils, Estación Experimental Agropecuaria, Instituto Nacional de Tecnologia y Agropecuaria, Sáenz Peña, Argentina.
1993-1999 - Editor, Coleopterists Bulletin.
 
Publications:
 
Clark, W. E. 1971. A taxonomic revision of the weevil genus Tychius Germar (Coleoptera: Curculionidae) in America north of Mexico. Brigham Young University Science Bulletin, Biological Series 13(3):1-28. 
Burke, H. R., W. E. Clark, and G. W. Frankie. 1975. Observations on the bionomics of Myrmex laevicollis (Horn) (Coleoptera: Curculionidae). Journal of the Kansas Entomological Society 48(2):160-169. 
Rogers, C. W., W. E. Clark, and H. R. Burke. 1975. Biosystematics of Sibinia sulcatula (Casey) on mesquite. The Southwestern Naturalist 20(3):303-314. 
Burke, H. R., and W. E. Clark. 1976. Cienfuegosia drummondii as a host of the boll weevil, Anthonomus grandis, in south Texas. Pages 12-26, in: Boll Weevil Suppression, Management, and Elimination Technology, Proceedings of a Conference, February 13-15, 1974, ARS-S71, USDA, ARS, 172 p. 
Clark, W. E. 1976. Review of the genus-group taxa included in the genus Tychius Germar (Coleoptera: Curculionidae). Entomologica Scandinavica 7:91-95. 
Clark, W. E. 1976. Notes on the life history and habits of Perigaster cretura (Herbst) (Coleoptera: Curculionidae) with descriptions of the larva and pupa. The Coleopterists Bulletin 31(2):159-165. 
Clark, W. E., G. Ibarra, and H. W. Van Cleave. 1976. Taxonomy and biology of spittlebugs of the genera Aeneolamia Fennah and Prosapia Fennah (Cercopidae) in northeastern Mexico. Folia Entomológica Mexicana 34:13-24. 
Clark, W. E., R. E. Warner, and D. R. Whitehead. 1977. Classification of the weevil subfamily Tychiinae, with a new genus and species, new combinations, and new synonymy in Lignyodini (Coleoptera: Curculionidae). The Coleopterists Bulletin 31(1):1-18. 
Clark, W. E. 1977. The male genital complex of some Curculionoidea (Coleoptera): description of associated musculature and discussion of function. The Coleopterists Bulletin 31(2):101-115. 
Clark, W. E. 1977. New synonymy, new combinations, a lectotype designation, and a type-species designation in Neotropical Tylodina (Coleoptera: Curculionidae, Cryptorhynchinae). Entomological News 88(7/8):199-201. 
Clark, W. E. 1977. North American Tychius: new synonymy and observations on phylogeny and zoogeography (Coleoptera: Curculionidae). Entomologica Scandinavica 8(4):287-300. 
Clark, W. E., and H. R. Burke. 1977. The curculionid genus Tychius Germar: natural history and coevolution with leguminous host plants. The Southwestern Entomologist 2(3):106-120. 
Clark, W. E.  1978.  The weevil genus Sibinia:  natural history, taxonomy, phylogeny, and zoogeography, with revision of the New World species.  Quaestiones Entomolgicae 14(2):91-387.
Clark, W. E. 1978. Revision of the Neotropical weevil genus Phymatophosus Faust (Coleoptera: Curculionidae). Systematic Entomology 3:103-130. 
Clark, W. E. 1978. Notes on the life history, and descriptions of the larva and pupa of Neotylopterus pallidus (LeConte) (Coleoptera: Curculionidae), a seed predator of Forestiera acuminata (Michx.) Poir. (Oleaceae). The Coleopterists Bulletin 32(2):177-184. 
Clark, W. E. 1978. The status of Paykull's Curculio fuscirostris (Coleoptera: Curculionidae). Entomologica Scandinavica 9:239-240. 
Clark, W. E., H. R. Burke, and D. M. Anderson. 1978. The genus Tychius Germar (Coleoptera: Curculionidae): larvae and pupae of some species, with evaluation of their characters in phylogenetic analysis. Proceedings of the Entomological Society of Washington 80(4):626-654. 
Clark, W. E. 1979. Taxonomy and biogeography of weevils of the genus Sibinia Germar (Coleoptera: Curculionidae) associated with Prosopis (Leguminosae: Mimosoideae) in Argentina. Proceedings of the Entomological Society of Washington 81(2):153-170. 
Clark, W. E. 1979. New species and new records of Sibinia Germar (Coleoptera: Curculionidae) from Panama. The Coleopterists Bulletin 33(2):209-216. 
Clark, W. E. 1979. A new species of Sibinia Germar from Mexico (Coleoptera: Curculionidae). The Coleopterists Bulletin 33(3):343-346. 
Clark, W. E. 1980. Revision of the weevil genus Neotylopterus Hustache (Coleoptera: Curculionidae). Annals of the Entomological Society of America 73(2):216-230. 
Clark, W. E. 1980. Revision of the weevil genus Lignyodes Dejean (Coleoptera: Curculionidae) in America north of Mexico. Transactions of the American Entomological Society 106(3):273-326. 
Clark, W. E. 1980. Additions to the Neotropical weevil genus Rosella Whitehead (Coleoptera: Curculionidae): R. triophori (Gyllenhal), n. comb., transferred from Lignyodes Dejean, and R. arcuata, n. sp. The Coleopterists Bulletin 34(3):299-304. 
Clark, W. E. 1981. New species and new records of Brazilian Sibinia Germar (Coleoptera: Curculionidae) associated with Mimosa scabrella Bentham (Leguminosae). The Coleopterists Bulletin 35(2):223-228. 
Clark, W. E. 1981. The weevil genus Hamaba Casey (Coleoptera: Curculionidae). The Florida Entomologist 64(2):217-222. 
Clark, W. E., and D. M. Anderson. 1981. The genus Chionanthobius Pierce (Coleoptera: Curculionidae): description of a new species on Forestiera (Oleaceae), and the larva and pupa of C. schwarzi Pierce. Proceedings of the Entomological Society of Washington 83(2):185-197. 
Clark, W. E. 1981. The genus Chionanthobius Pierce (Coleoptera: Curculionidae): a new species from Cuba and a key to species. Proceedings of the Entomological Society of Washington 83(4):690-693. 
Clark, W. E., and N. Lodos. 1981. Notes on Turkish Lignyodes Dejean (Coleoptera: Curculionidae), with description of a new species. The Coleopterists Bulletin 35(3):311-315. 
  
Clark, W. E.  1982.  Classification of the weevil tribe Lignyodini (Coleoptera:  Curculionidae, Tychiinae), with revision of the genus Plocetes.  Transactions of the American Entomological Society 108(1&2):11-151. 
Clark, W. E. 1982. Systematic studies on weevils in the family Curculionidae. Highlights of Agricultural Research (Alabama Argicultural Experiment Station) 29(3):6. 
Clark, W. E. 1982. Apolpones Hustache a new synonym of Sibinia Germar (Coleoptera: Curculionidae). The Coleopterists Bulletin 36(2):287-290. 
Clark, W. E. 1983. New species and new records of the lignyodine weevil genus Plocetes LeConte (Coleoptera: Curculionidae) from Chiapas, Mexico. Proceedings of the Entomological Society of Washington 85(4):797-805.
Burke, H. R., W. E. Clark, and W. H. Cross. 1984. Larvae and pupae of the Anthonomus subgenus Anthonomorphus Dietz, A. grandis Boheman and A. hunteri Burke and Cate (Coleoptera: Curculionidae). The Southwestern Entomologist 9(1):84-90.
Clark, W. E. 1984. Curculio picirostris Fabricius, 1787, and Tychius stephensi Schönherr, 1836 (Coleoptera: Curculionidae): proposed conservation under the plenary powers. Z.N.(S.)2266. Bulletin of Zoological Nomenclature 41(1):45-52. 
Clark, W. E. 1984. Species of Sibinia Germar (Coleoptera: Curculionidae) associated with Mimosa pigra L. Proceedings of the Entomological Society of Washington 86(2):358-368.
Clark, W. E. 1984. A new Mexican Sibinia Germar allied to S. bothrosterna Clark (Coleoptera: Curculionidae). The Coleopterists Bulletin 38(4):390-393.
Clark, W. E., and H. R. Burke. 1985. Revision of the venustus species group of the weevil genus Anthonomus Germar (Coleoptera: Curculionidae). Transactions of the American Entomological Society 111(1):103-170.
Clark, W. E., and H. R. Burke. 1986. Revision of the gularis species group of the genus Anthonomus Germar (Coleoptera: Curculionidae). The Coleopterists Bulletin 40(1):1-26.
Clark, W. E., and H. R. Burke. 1986. A new Neotropical species of Anthonomus Germar (Coleoptera: Curculionidae) associated with Bombacopsis quinata (Bombacaceae). Proceedings of the Entomological Society of Washington 88(2):320-327.
Clark, W. E., and H. R. Burke. 1986. Phylogeny of the species of the Anthonomus subgenus Anthonomorphus Dietz, with discussion of relationships with Anthonomus grandis Boheman (Coleoptera: Curculionidae). Journal of the Kansas Entomological Society 59(3):508-516.
Burke, H. R., W. E. Clark, J. R. Cate and P. A. Fryxell. 1986. Origin and dispersal of the boll weevil. Bulletin of the Entomological Society of America 32(4):228-238.
Clark, W. E. 1987. Revision of the unipustulatus group of the weevil genus Anthonomus Germar (Coleoptera: Curculionidae). The Coleopterists Bulletin 41(1):73-88.
 
Clark, W. E.  1987.  Revision of the Nearctic species of Pseudanthonomus Dietz (Coleoptera: Curculionidae).  The Coleopterists Bulletin 41(3):263-285.
Clark, W. E. 1987. Revision of the Nearctic species of Pseudanthonomus Dietz (Coleoptera: Curculionidae). The Coleopterists Bulletin 41(3):263-285. 
Clark, W. E., and R. P. Martíns. 1987. Anthonomus biplagiatus Redtenbacher (Coleoptera: Curculionidae), a Brazilian weevil associated with Kielmeyera (Guttiferae). The Coleopterists Bulletin 41(2):157-164.
Clark, W. E. 1987. Revision of the Anthonomus subgenus Anthomorphus Weise (Coleoptera: Curculionidae). Quaestiones Entomologicae 23:317-364. 
Clark, W. E. 1987. The species of Anthonomus in the albolineatus group (Coleoptera: Curculionidae). Transactions of the American Entomological Society 113(4):309-359. 
Clark, W. E. 1988. Revision of the furcatus species group of the weevil genus Anthonomus Germar (Coleoptera: Curculionidae). The Coleopterists Bulletin 42(4):359-377. 
Clark, W. E. 1988. Review of the Anthonomus alboscutellatus species group, with description of a new species (Coleoptera: Curculionidae). The Coleopterists Bulletin 42(4):379-386. 
  
Clark, W. E.  1988.  Revision of the weevil genus Loncophorus Chevrolat (Coleoptera: Curculionidae, Anthonominae).  Quaestiones Entomologicae 24(3):465-518.
Clark, W. E., and H. R. Burke. 1989. Revision of the ornatus species group of the genus Anthonomus Germar (Coleoptera: Curculionidae, Anthonominae). Proceedings of the Entomological Society of Washington 91(1):88-111. 
Clark, W. E. 1989. Revision of the puncticeps species group of the genus Anthonomus Germar (Coleoptera: Curculionidae). The Coleopterists Bulletin 43(1):45-57. 
Clark, W. E. 1989. Lonchophorellus Clark, a New Genus of Neotropical Anthonomini (Coleoptera: Curculionidae). The Coleopterists Bulletin 43(3):279-289. 
Clark, W. E. 1989. Revision of the Neotropical Weevil Genus Atractomerus Duponchel and Chevrolat (Coleoptera: Curculionidae). Transactions of the American Entomological Society 114(4):313-413. 
Clark, W. E. 1990. Revision of the flavirostris species group of the genus Anthonomus Germar (Coleoptera: Curculionidae). Transactions of the American Entomological Society 116(1):261-294. 
Clark, W. E. 1990. Revision of the fischeri species group of the genus Anthonomus Germar (Coleoptera: Curculionidae). Transactions of the American Entomological Society 116(3):619-642. 
Clark, W. E. 1990. Revision of the calvescens species group of the genus Anthonomus Germar (Coleoptera: Curculionidae). Transactions of the American Entomological Society 116(3):643-654. 
Clark, W. E. 1990. The Neotropical species of Pseudanthonomus Dietz (Coleoptera: Curculionidae). Transactions of the American Entomological Society 116(3):655-695. 
Clark, W. E. 1990. Revision of the Anthonomus subgenus Anthonomocyllus Dietz (Coleoptera: Curculionidae). Quaestiones Entomolgicae 26 (4):559-600. 
Clark, W. E. 1991. The Anthonomus guttatus species group (Coleoptera: Curculionidae). Proceedings of the Entomological Society of Washington 93(2):262-271. 
Clark, W. E. 1991. Dietzia Champion, a new synonym of Plocetes LeConte (Coleoptera: Curculionidae). The Coleopterists Bulletin 45(2):191-192. 
Clark, W. E. 1991. Revision of the Anthonomus alboannulatus and Anthonomus triensis species groups (Coleoptera: Curculionidae). Coleopterists Bulletin 45(3):206-226. 
Clark, W. E. 1991. The Anthonomus curvirostris species group Transactions of the American Entomological Society 117(1):39-66. 
Clark, W. E. 1991. The Anthonomus rubiginosus species group (Coleoptera: Curculionidae). Transactions of the American Entomological Society 117(3-4):145-166. 
Clark, W. E. 1992. The Anthonomus partiarius species group (Coleoptera: Curculionidae). Transactions of the American Entomological Society 118(1):107-128. 
Clark, W. E. 1992. The Anthonomus marmoratus species group (Coleoptera: Curculionidae). Transactions of the American Entomological Society 118(1):129-145. 
Clark, W. E. 1992. The Biologia Centrali-Americana species of Anthonomus in the subgenus Anthomorphus Weise (Coleoptera: Curculionidae). The Coleopterists Bulletin 46(2):203-207. 
Clark, W. E. 1992. New synonymy, a new combination and a lectotype designation in neotropical Anthonomini (Coleoptera: Curculionidae). The Coleopterists Bulletin 46(3):286-289. 
Clark, W. E. 1993. The weevil genus Neomastix Dietz (Coleoptera: Curculionidae). The Coleopterists Bulletin 47(1):1-19. 
Clark, W. E. 1993. The Anthonomus cavei species group (Coleoptera: Curculionidae). Proceedings of the Entomological Society of Washington 95(2):152-162. 
Clark, W. E. 1993. Huaca Clark, a new genus of neotropical Anthonomini (Coleoptera: Curculionidae). Transactions of the American Entomological Society 119(1):1-45. 
Clark, W. E. 1993. The Anthonomus ocularis species group (Coleoptera: Curculionidae).  Transactions of the American Entomological Society 119(2):95-130. 
Clark, W. E. 1993. Listrorrhynchus Champion, a new synonym of Anthonomus Germar (Coleoptera: Curculionidae). Coleopterists Bulletin. 47(2):192-194. 
Clark, W. E. 1993. The Anthonomus monostigma species group (Coleoptera: Curculionidae). Transactions of the American Entomological Society 119(4):231-245. 
Clark, W. E. 1993. The Anthonomus ourateae, A. planipennis and A. albocivitensis species groups, and new species in the A. rubiginosus and A. partiarius groups (Coleoptera: Curculionidae).
Transactions of the American Entomological Society 119(4):247-269. 
Clark, W. E. 1994. The Anthonomus gibbicrus species group (Coleoptera: Curculionidae). Proceedings of the Entomological Society of Washington 96(2)193-198. 
Clark, W. E. 1994. New species of Anthonomus Germar in the A. dogma and A. furcatus species groups (Coleoptera: Curculionidae). Proceedings of the Entomological Society of Washington 96(2):199-207. 
Clark, W. E. 1995. A new species of Loncophorus from Mexico (Coleoptera: Curculionidae, Anthonomini) associated with Mortoniodendron (Malvales: Tiliaceae). Anales del Instituto de Biología de la Universidad Nacional Autónima de México, Serie Zoologica 66(1):107-112. 
Clark, W. E. 1996. New species in the Anthonomus salvini group (Coleoptera: Curculionidae: Anthonomini). Proceedings of the Entomological Society of Washington 98(2):267-273. 
  
Clark, W. E., and H. R. Burke.   1996.  The species of Anthonomus Germar (Coleoptera: Curculionidae) associated with plants in the family Solanaceae.  Southwestern Entomologist.  Supplement, 19 Southwestern Entomological Society, College Station, TX. 114p.
Revision of the Weevil Genera Magdalinops Dietz and Chelonychus Dietz (Coleoptera: Curculionidae, Anthonomini). The Coleopterists Bulletin 56(1):107–122.
Clark, W. E., and H. R. Burke.   2001.  Revision of the weevil genus Epimechus Dietz (Coleoptera: Curculionidae: Anthonomini).
Insecta Mundi.  15(2):95-116.
Clark, W.E., and H.R. Burke. 2005. Revision of the subgenus Cnemocyllus Diets of the weevil genus Anthonomus Germar (Coleoptera: Curculionidae, Anthonomini). Insecta Mundi 19(1-2)1-54. 
  Wayne E. Clark's Personal Page 
 
 
 
 
 The Insect to the left
 is a paratype of Sibinia fastigiata Clark (Coleoptera: Curculionidae)
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