| Bulletin 441
December 1972
Alabama Agricultural Experiment Station
R. Dennis Rouse,
Director
Auburn University
Auburn, Alabama |
Genus
Capraita Bechyné
Capraita Bechyné, 1957,
Ann. Mus. Civ. Storia Nat. Giacoma Doria 69: 51-74.
Chloephaga Weise, 1899, Deutsche Entomol. Zeitschr. 1899: 138
(not Eyton, 1838).
Small (3.0-6.0 mm.), depressed. Antennae
slender, at least 1/2 body length. Eyes often large and closely set. Lateral
margins usually explanate. Prothorax at base nearly as wide as elytra.
Elytra confusedly punctate or impunctate and glabrous. Prosternum wide
between anterior coxae, at least 1/2 coxal width. Apical hind tarsal segment
globose, basal joint comparatively short.
In placing the following species in Capraita,
we are following Bechyné (11), who first separated the
species of Oedionychus Series B (52) from the true Oedionychus
(apterous Mediterranean forms). At the time of this split (1955), these
North American species were placed in Chloephaga Weise, which
name has been proven to be a junior generic homonym. The new name, Capraita
Bechyné (12) replaced Chloephaga Weise.
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| KEY
TO CAPRAITA |
1. Elytra with inconspicuous umbones
and basal callosities; explanate margins narrow….. 2
Elytra with distinct umbones and basal callosities; explanate margins
wide….. 3
2. Elytra entirely dark blue-black (Fig. 31)….. C. indigoptera
Elytra black with margins yellow or yellow with large discal black spot…..
C. thyamoides
3. Dorsal profile broadly oval, sides of elytra arcuate….. 4
Dorsal profile elongate, sides of elytra subparallel….. 5
4. Elytra dark brown-black with apex broadly and lateral margins narrowly
light yellow -brown….. C. saltatra
Elytra entirely dark brown-black….. C. quercata
5. Head coarsely punctate….. 6
Head faintly punctate, nearly smooth….. 7
6. Pronotum coarsely punctate, elytra irregularly fasciate and maculate…..
C. sexmaculata
Pronotum impunctate or only with very fine punctures, elytra with red
sinuate sutural vitta or rarely entirely
red with faint yellow spots….. C. suturalis
7. Elytra with prominent basal callosities; eyes large, interocular
space subequal to 1/3 width of head…..
8
Elytra lacking basal callosities; flatter, less convex; eyes small,
interocualr space subequal to 1/2 width
of head….. C. circumdata
8. Small (length 3.5-4.7 mm.), elytral punctation fine….. 9
Large (length 4.7-5.6 mm.), elytral punctation coarse….. C.
scalaris
9. Pronotum and elytra dark brown or black, lateral margins pale…..
C. obsidiana obsidiana
Pronotum and elytra yellow marked with dark brown or black fasciae and
spots, not entirely dark with light margins…..
C. obsidiana blakeae
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| C.
INDIGOPTERA (LECONTE) |
Oedionychis indigopthera LeConte, 1878,
Proc. Amer. Philos. Soc. 17: 416.
Elongate oval. Head and pronotum red-brown;
elytra blue-black. Head alutaceous, finely sparsely punctate. Antennal
segments 1 thorugh 4 smoother dark red-brown, 5-11 finely reticulate,
black. Pronotum alutaceous, very finely punctate. Elytra alutaceous,
densely punctate, not very convex, entirely blue-black; humeri only
slightly prominent with slight interhumeral depressions. Length 3.5-4.9
mm. Width 1.9-2.5 mm.
Alabama records: 21 specimens from Lee1,
Macon1, and Mobile2,3 counties.
Seasonal distribution: April 7-September
13.
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| C.
THYAMOIDES (CROTCH)* |
Capraita
thyamoides (Crotch)*
Oedionychis thyamoides Crotch,
1873, Proc. Acad. Nat. Sci. Philadelphia 25: 63.
Oval. Head and prothorax yellow brown,
elytra black with yellow margin or variable width. Head alutaceous,
relatively coarsely but sparsely punctate. Prothorax alutaceous, finely
punctate. Elytra alutaceous, densely, strongly punctate; margin moderately
explanate. Length 4.2-4.6 mm. Width 2.3-2.5 mm.
Alabama records: 2 specimens from Marion1
and Marshall1 counties.
Seasonal distribution: July 12-28.
Remarks: Blake (15) recorded
Teucrium canadense as host plant.
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| C.
SALTATRA (BLATCHLEY) |
Capraita
saltatra (Blatchley)
Oedionychis saltatra Blatchley,
1923, Canadian Entomol. 55: 32.
Oval. Head and prothorax yellow-brown.
Elytra piceous with lateral margins near humeri narrowly yellow and
apexes yellow or dirty brown (lighter than disc). Surface shining, not
at all or slightly alutaceous. Length 4.2-4.9 mm. Width 2.5-3.0 mm.
Alabama records: 6 specimens from Lee,
Mobile2,3, and Russell counties.
Seasonal distribution: January 14, March
16-August 10.
Remarks: These beetles have been taken
January 14 under bark (75). It is possible that comparisons
of the aedeagi will further indicate the conspecificity of this and
C. quercata (Fabricius). There are specimens in Loding's collection
from Chastang, Mobile County, January 14, which are intermediate between
these two species.
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| C.
QUERCATA (FABRICIUS) |
Capraita
quercata (Fabricius)
Galleruca quercata Fabricius,
1801, Syst. Eleuth. I., p. 495.
Oval. Head and prothorax yellow-brown.
Elytra entirely piceous with lateral margins near humeri slightly ferruginous.
Surface minutely alutaceous. Length 4.2-5.4 mm. Width 2.5-3.3 mm.
Alabama records: 8 specimens from Baldwin1
and Mobile1,2,3 counties.
Seasonal distribution: January 1-July.
Remarks: Capraita saltatra may
be found to be conspecific with C. quercata.
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| C.
SEXMACULATA (ILLIGER) |
Capraita
sexmaculata (Illiger)
Haltica sexmaculata Illiger,
1807, Mag. Insektenk. 6: 104.
Haltica palliata Randall, 1838, Boston Jour. Nat. Hist. 2:
47.
Chloephaga creticus Jacoby, 1886, Entomol. Zeit. 47: 215.
Oblong. Head alutaceous, clypeus and frons
yellow-brown, vertex dark brown, punctate with wide punctures. Pronotum
strongly alutaceous, coarsely punctate, light yellow to tan laterally
and midbasally, remainder of disc with brown to black irregular maculae.
Elytra densely, coarsely punctate with submarginal plica; yellow with
piceous maculae: humeral spots, basal medial cordate spot, confluent
mid- and subapical lateral spots, and elongate broad medial subapical
spots, latter occasionally as broad fascia. Length 3.2-4.0 mm. Width
1.8-2.1 mm.
Alabama records: 16 specimens from Baldwin3,
Clay1, DeKalb1, Houston1, Lee1,
Mobile2,3, and Tallapoosa1 counties.
Seasonal distribution: January 9, April
25-June 25.
Remarks: Capraita sexmaculata
has been reported from on Fraxinus sp. and Chionanthus
virginica (15).
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| C.
SUTURALIS (FABRICIUS) |
Capraita
suturalis (Fabricius)
Galleruca suturalis Fabricius,
1801, Syst. Eleuth. I., p. 499.
Oblong oval. Frons and vertex usually
red-brown but in some specimens yellow. Pronotum yellow. Elytra yellow
or yellow (entirely so in some samples), but usually with midbase, scutellar
area, and suture red, or rarely varying to nearly entirely red with
only faint subapical yellow spots and lateral margins lighter basally.
Length 3.8-4.4 mm. Width 2.1-2.5 mm.
Alabama records: 19 specimens from Baldwin1
and Mobile2,3 counties.
Seasonal distribution: March 31-August
27.
Remarks: We have collected this species
in Baldwin County both in the day and at night by beating plants of
the Family Ericaceae.
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| C.
CIRCUMDATA (RANDALL) |
Capraita
circumdata (Randall)
Haltica circumdata Randall,
1838, Boston Jour. Nat. Hist. 2: 48.
Aedionychis limbalis Melsheimer, 1847, Proc. Acad. Nat. Sci.
Philadelphia 3: 162.
Oedionychis quercata Crotch, 1873, Proc. Acad. Nat. Sci. Philadelphia
25: 63 (not Fabricius).
Oblong oval. Head, prothorax, and mesothorax
ventrally, yellow-brown; occiput slightly darker. Elytra black, margined
with yellow, margin slightly wider at apex, rarely with two small oblique
yellow spots at basal 1/3; somewhat shining, sides subparallel, margin
explanate; umbone well defined with depression between it and basomedial
callosity. Metasternum and abdomen black. Length 3.9-4.9 mm. Width 2.1-2.8
mm.
Alabama records: 80 specimens from DeKalb1,
Etowah1, Lee1, Macon1, Madison1,
Mobile2, Tallapoosa1, and Winston1,2
counties.
Seasonal distribution: March 27-August
3.
Remarks: We have most frequently taken
these beetles by beating Vaccinum spp.; however, Blake (15)
has taken them on Fagus grandifolia, Juglans cinerea, Plantago
lanceolata, and Verbena urticifolia. We also collected
a single specimen on Rubus sp. upon which it may have been
only resting.
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| C.
OBSIDIANA OBSIDIANA (FABRICIUS) |
Capraita
obsidiana obsidiana (Fabricius)
Galleruca obsidiana Fabricius,
1801, Syst. Eleuth., I., p. 499.
Oedionychis quercata Horn, 1889, Trans. Amer. Entomol. Soc.
16: 194 (not Fabricius).
Oblong oval. Labrum and clypeus yellow,
frons and occiput black, alutaceous, punctate. Pronotum alutaceous,
finely punctate, lateral margins yellow. Elytra black, basally alutaceous,
punctate; lateral margins yellow to basal 1/3, apexes yellow forming
an arcuate emargination into black color of disc. Length 3.5-4.2 mm.
Width 2.0-2.5 mm.
Alabama records: 24 specimens from Barbour2,
Houston1, and Mobile3 counties.
Seasonal distribution: April 25-August
10.
Remarks: Nearly all of the Houston County
specimens were taken by beating Vaccinium sp.
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| C. OBSIDIANA
BLAKEAE (LENG AND MUTCHLER) |
Capraita
obsidiana blakeae (Leng and Mutchler)
Oedionychis obsidiana flava
Blake, 1927, Proc. U. S. Nat. Mus. 70(23): 40 (var.) (not Gmelin).
Oedionychis obsidiana blakeae Leng and Mutchler, 1933, 2nd and
3rd Supp. to Cat. of Coleopt. of America
N. of Mexico, p. 45.
Oblong oval. Labrum and clypeus yellow,
frons and occiput dark brown to piceous, alutaceous, punctate. Pronotum
alutaceous, very finely punctulate, yellow with two brown discal spots,
spots frequently confluent. Elytra yellow, punctate with black maculae
of various configurations; sometimes basal or subapical fasciae, or
lateral spots joined in tandem. Length 3.9-4.7 mm. Width 1.9-2.5 mm.
Alabama records: 44 specimens from Baldwin1,
Barbour2, Clay1, Lee1, Lowndes1,
Macon1, Marengo1, Mobile2, and Talladega1
counties.
Seasonal distribution: March 27-August
27.
Remarks: This subspecies has also been
collected on Vaccinium sp. Specimens intermediate between the
lighter forms of C. obsidiana blakeae and the nominate subspecies
have been collected in Baldwin County.
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| C.
SCALARIS (MELSHEIMER) |
Capraita
scalaris (Melsheimer)
Oedionychis scalaris Melsheimer,
1847, Proc. Acad. Nat. Sci. Philadelphia 3: 163.
Oedionychis lobata LeConte, 1859, Coleopt. Kans., p. 24.
Broadly oblong oval. Head yellow, sometimes
with brown spot on occiput, eyes large and closely set. Pronotum 2 1/2
times as wide as long, yellow, with broad explanate margin. Elytra shining,
rather strongly, closely punctate; umbones distinct with depression
between them and basomedial callosities; yellow with black markings:
usually suture irregularly vittate to apical declivity, often with postbasal,
midelytral, and preapical fasciae. Length 4.7-5.6 mm. Width 2.5-2.8
mm.
Alabama records: 5 specimens from Lee1,
Macon1, and Mobile2 counties.
Remarks: Blatchley, according to Blake
(15), collected this species in Florida on ericaceous plants.
Since there is some difficulty encountered in separating specimens of
this species from those of C. obsidiana blakeae, aedeagi were
extracted and compared. The results of the comparison indicated the
specfic distinctness of these forms. "It is to be separated from
the banded forms of obsidiana var. flava (synonym
of blakeae L. & M.) by its relatively larger size and coarser
punctation" (15).
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